Kenneth B. Armitage Ecology and Evolutionary Biology Department, The University of Kansas marmots@ukans.edu 1142364203256 These data continue and expand upon data collected by Kenneth B. Armitage titled, " Social Behavior and Population Dynamics of Yellow-bellied Marmots", that goes from 1965 - 2001. A record includes the subject's age, sex, body mass, locality, and reproductive status. Starting in 2002, additional columns are added that include: the subject's reaction to being trapped and tagged, and a list of samples that were collected (blood, feces, hair, alarm call recordings). To access data from 2002, please contact Daniel Blumstein. Marmot Armitage yellow-bellied marmot Marmota flaviventris Rocky Mountain Biological Laboratory mammal mass Rodentia Alpine/Tundra Population Dynamics Life History Community Structure Survival Please contact Daniel Blumstein regarding access to the data. Rocky Mountain Biological Laboratory -106.989 -106.989 38.959 38.959 Species Marmota flaviventris Genus Marmota 1965-06-11 1142363819896 Dan Jones Rocky Mountain Biological Laboratory

P.O. Box 519 Crested Butte CO 81224

970-349-7231 it@rmbl.org Daniel Blumstein UCLA, Department of Ecology and Evolutionary Biology 310-267-4746 marmots@ucla.edu Janice Daniel The Blumstein Lab, UCLA Marmot database manager janice@ucla.edu

Marmots were live-trapped and age, sex, reproductive condition, and body mass were recorded. Each animal received a unique, four-digit ear tag in its left and right ear. Selected animals were marked with non-toxic fur dye for individual recognition during behavioral observations from afar.

public read uid=janice,o=unaffiliated,dc=ecoinformatics,dc=org read write cn=knb-prod,o=NCEAS,dc=ecoinformatics,dc=org read write Marmot Trap Log 1965 and Continuing Marmots were live-trapped and age, sex, reproductive condition, and body mass were recorded. Each animal received a unique, numbered ear tag in its left and right ear. Selected animals were marked with non-toxic fur dye for individual recognition during behavioral observations from afar. Trap _Log_1965_continue.txt 1 #x0A column #x09 request data from contact Janice_ notes Janice's notes Janice Daniel's notes notes Year Year data collected YYYY 1 Location Location description through 1996 only D down valley M mid valley U up valley GROUP Group Code (through 1996 only) F/00 = young female (pup) F/01 = yearling female (1 year old) F/02/NR = non-reproductive 2 year old female F/NR = non-reproductive adult female F/RR = reproductive female M/00 = young male (pup) M/01 = yearling male (1 year old) M/AA = adult male F/00 young female (pup) F/01 yearling female (1 year old) F/02/NR non-reproductive 2 year old female F/NR non-reproductive adult female F/RR reproductive female M/00 young male (pup) M/01 yearling male (1 year old) M/AA adult male Colony_Number Colony # Colony # 1 = River colony, 2 = Bench colony, 3 = Gothic talus colony, 4 = Marmot meadow colony, 50 = All picnic colony, 51 = Upper picnic colony, 52 = Middle picnic colony, 53 = Lower picnic colony, 6 = Boulder colony, 7 = North picnic colony, 8 = Cliff colony, 9 = Bellview talus colony, 10 = Grasshopper ledge colony location. 1 River colony 2 Bench colony 3 Gothic talus colony 4 Marmot meadow colony 50 All picnic colony 51 Upper picnic colony 52 Middle picnic colony 53 Lower picnic colony 6 Boulder colony 7 North picnic colony 8 Cliff colony 9 Bellview talus colony 10 Grasshopper ledge colony RColony Colony location, Many of these location names have photos that contain a diagram of the location. (These jpg files are included in this dataset.) Here are some colony corrdinates: River - at the Spruce Tree 2867 m N 38degrees 56.947', W 106degrees 59.270' ; Marmot Meadow - at Main Talus 2930m N 38degrees 58.751', W 106degrees 59.999' ; North Picnic - at Slope Burrow 3008m N 38degrees 59.203', W 107degrees 00.791' ; Boulder - at the big boulder 2971m N 38 degrees 59.125', W 107degrees 00.660' ; Picnic - at Split Rock 2992m N 38 degrees 59.011', W 107degrees 00.664' Bellview BelowBellview.JPG Bench Bench.JPG , BelowBellview.JPG Corral burrow RiverCoralBurrow.JPG Free text (unrestricted) .* Burrow Burrow location, Many of these location names have photos that contain a diagram of the location. (These jpg files are included in this dataset.) Bench Cabin BenchCabinAndTowerBurrow.JPG L_tag L tag Left ear tag number. Each subject is given a set of left and right ear tags when first trapped (original tags). These tags are sometimes lost and replaced with different numbered tags over time. The 'Ltag' and 'Rtag' columns came from the original trap log, and sometimes represent(ed) the tags as they exist(ed) on the marmots themselves at the time of trapping, but other times give the original tags. In rare instances, different subjects were given identical combinations of left and right tags. A suffix of 'a', 'b', or 'c' was added to the right tag in cases of identical tag combinations. Other changes to the tags should be noted in the 'Janice's notes' column. ID number R_tag R tag Right ear tag number. Each subject is given a set of left and right ear tags when first trapped (original tags). These tags are sometimes lost and replaced with different numbered tags over time. The 'Ltag' and 'Rtag' columns came from the original trap log, and sometimes represent(ed) the tags as they exist(ed) on the marmots themselves at the time of trapping, but other times give the original tags. In rare instances, different subjects were given identical combinations of left and right tags. A suffix of 'a', 'b', or 'c' was added to the right tag in cases of identical tag combinations. Other changes to the tags should be noted in the 'Janice's notes' column. ID number database_unique_ID database unique ID Unique ID for record in database. Comprised of original left and right ear tag numbers. Suffixes 'a', 'b', and 'c' added when multiple subjects shared identical original tags. ID number current_left current left Left ear tag on subject at time of trapping. May or may not be the same as the original ear tag, depending upon whether ear tags were lost and replaced over time. Entries for 1996-1999 only. ID number current_right current right Right ear tag on subject at time of trapping. May or may not be the same as the original ear tag, depending upon whether ear tags were lost and replaced over time. Entries for 1996-1999 only. ID number sex Subject's sex, either male or female. sex AG_mm AG mm Ano-genital distance (distance between center of anus and genital pore) in millimeters millimeter 0.1 real Age Age Marmot Age - Note that 21+ means that the individual was an adult immigrant to the site, with 21 representing the first year seen, 22 the second year seen, etc. When individuals are trapped as pups, then their exact age will be known from year to year. 0 = juvenile (pup) - born in the summer of that year, usually around late June-July. 1 = yearling - one year old - born previous summer. 2 = 2 years old. 15 = 15 years old - oldest recorded female. An individual that was first recorded as an adult is given a series of numbers starting with 21. At age 21, the individual is at least 2 years old. No individual has ever lived as long as 21 years. 21 = immigrant or previously untrapped individual of unknown age, at least 2 years old. 1st season recorded. 22 = immigrant or previously untrapped individual of unknown age, at least 3 years old. 2nd season recorded. 23 = 3rd season recorded. At least 4 years old. An individual first recorded as being of unknown age (but is clearly not a pup by weight) is given a series of numbers starting with 61. At age 61, the individual is at least 1 years old. 61 = immigrant or previously untrapped individual, most likely a yearling, but could be a small adult, at least 1 year old. 1st season recorded. 62 = 2nd season recorded, now at least 2 years old. 63 = 3rd season recorded. At least 3 years old. This numbering system is not consistent throughout. 99 means of unknown age. A means adult of unknown age. 0 juvenile (pup) - born in the summer of that year, usually around late June-July. 1 yearling - one year old - born previous summer. 2 2 years old 15 15 years old - oldest recorded female. 21 immigrant or previously untrapped individual of unknown age, at least 2 years old. 1st season recorded. 22 immigrant or previously untrapped individual of unknown age, at least 3 years old. 2nd season recorded. 23 3rd season recorded. At least 4 years old. 61 immigrant or previously untrapped individual, most likely a yearling, but could be a small adult, at least 1 year old. 1st season recorded. 62 2nd season recorded, now at least 2 years old. 63 3rd season recorded. At least 3 years old. 99 unknown age A adult of unknown age R_age R age The R_age includes corrections or additions to the Age column, with R2_Age giving the final version of the age. (note from Janice Daniel) 0 juvenile (pup) - born in the summer of that year, usually around late June-July. 1 yearling - one year old - born previous summer. 2 2 years old 15 15 years old - oldest recorded female. 21 immigrant or previously untrapped individual of unknown age, at least 2 years old. 1st season recorded. 22 immigrant or previously untrapped individual of unknown age, at least 3 years old. 2nd season recorded. 23 3rd season recorded. At least 4 years old. 61 immigrant or previously untrapped individual, most likely a yearling, but could be a small adult, at least 1 year old. 1st season recorded. 62 2nd season recorded, now at least 2 years old. 63 3rd season recorded. At least 3 years old. 99 unknown age A adult of unknown age R2_Age R2 Age The R_age includes corrections or additions to the Age column, with R2_Age giving the final version of the age. (note from Janice Daniel) 0 juvenile (pup) - born in the summer of that year, usually around late June-July. 1 yearling - one year old - born previous summer. 2 2 years old 15 15 years old - oldest recorded female. 21 immigrant or previously untrapped individual of unknown age, at least 2 years old. 1st season recorded. 22 immigrant or previously untrapped individual of unknown age, at least 3 years old. 2nd season recorded. 23 3rd season recorded. At least 4 years old. 61 immigrant or previously untrapped individual, most likely a yearling, but could be a small adult, at least 1 year old. 1st season recorded. 62 2nd season recorded, now at least 2 years old. 63 3rd season recorded. At least 3 years old. 99 unknown age A adult of unknown age s_st s st Unknown what this column refers to. Entries from 1979-1993. ID number Reproductive_Status Reproductive Status Reproductive Status - 1 = testes scrotal, 2 = testes unknown, 3 = nipples visible, 4 = nipples prominent, 5 = nipples swollen, 6 = lactating 1 testes scrotal 2 testes unknown 3 nipples visible 4 nipples prominent 5 nipples swollen 6 lactating Litter_Size Litter Size Litter Size Litter number Date Date Date MM-DD-YYYY 1 Time Time Time hh:mm 1minute Mass Weight of subject in grams. gram 1 natural Mark Mark refers to shape and location of mark on animal. Fur mark is applied using dye when animal is trapped. Used to visually identify individuals in the field. Field notebooks often reference fur marks, but then this info is converted to unique IDs in spreadsheets. Symbols hindfoot length of hindfoot in millimeters millimeter .01 real notes notes regarding individual marmots notes Tag_changes Changes to ear tags when lost and replaced. tag changes Defecated Defecated? Defecated: 0 = no, 1 = yes if the marmot defecated in the trap, 2 = yes if the marmot defecated in the handling bag, 3 = the marmot defecated in the trap and in the bag. To remove the subject from the trap, we place a handling bag over the trap entrance and allow the marmot to walk out of the trap into the bag. Ear tags and fur dye are applied, and other measurements and samples are taken while the subject is in the bag, prior to release. 0 no 1 yes if the marmot deficated in the trap 2 yes if the marmot defecated in the bag 3 the marmot defecated in the trap and in the bag Tooth_Chattered Tooth Chattered? Tooth chattered: 0 = no, 1 = yes if the marmot tooth chattered in the trap (chattering sound made by rapidly clacking upper and lower teeth together). 0 no 1 yes if the marmot tooth chattered in the trap Called Called? Called: 0 = no, 1 = yes if the marmot alarm called in the trap 0 no 1 yes if the marmot alarm called in the trap Struggled Struggled? Struggled in trap: 0 = no, 1 = yes if the marmot was struggling or banging itself against the trap. 0 no 1 yes Tried_to_bite? Tried to bite? Tried to bite through the cage: 0 = no, 1 = yes if the marmot was trying to bite through the cage. 0 no 1 yes Immed_Walked_into_bag Immed Walked into bag? Immediately walked into the bag: 0 = no, 1 = yes if the marmot walked from the trap into the handling bag more-or-less immediately. 0 no 1 yes Samples Samples taken: b = blood sample, f = fecal sample, h = hair, rec = recorded, run = run, v = vaginal cell sample b blood sample f fecal sample h hair rec recorded run run v vaginal cell sample Blood Blood? Blood sample taken: 0 = no, 1 = yes 0 no 1 yes Fecal Fecal? Fecal sample collected: 0 = no, 1 = yes 0 no 1 yes Hair Hair? Hair sample collected: 0 = no, 1 = yes 0 no 1 yes Recorded Recorded? 1 = subject was recorded alarm calling in trap, 0 = subject was not recorded. Applies to 2001 only. alarm call recorded Run Run? Maximum running speed measured: 0 = no, 1 = yes 0 no 1 yes BelowBellview.JPG Below Bellview Bench Parking Lot - photo includes Rocky Mountian Biological Laboratory (RMBL) Marmot trapping areas. This is part of the PowerPoint Slidshow MarmotTrappingAreasRMBL092005.ppt BelowBellview.JPG jpg ecogrid://knb/bowdish.155.1 BelowBellview Below Bellview Below Bellview Bench Parking Lot trapping area Stonefield_SouthMound.JPG Stonefield-South Mound - photo includes Rocky Mountian Biological Laboratory (RMBL) Marmot trapping areas. This is part of the PowerPoint Slidshow MarmotTrappingAreasRMBL092005.ppt Stonefield_SouthMound.JPG jpg ecogrid://knb/bowdish.156.1 StonefieldSouthMound Stonefield-South Mound Stonefield-South Mound trapping location Stonefield.JPG Stonefield - photo includes Rocky Mountian Biological Laboratory (RMBL) Marmot trapping areas. This is part of the PowerPoint Slidshow MarmotTrappingAreasRMBL092005.ppt . Includes: The Diamond, Turkey, Pebbles, Point, Fred, Rubble, Main Mound, Belly, Button, Platform, Valley, RockIsland, Post, Shark, Bam Bam, Pompei and South Mound locations Stonefield.JPG jpg ecogrid://knb/bowdish.157.1 Stonefield.JPG Stonefield marmot tagging areas tagging areas NorthPicnic.JPG North Picnic - photo includes Rocky Mountian Biological Laboratory (RMBL) Marmot trapping areas. This is part of the PowerPoint Slidshow MarmotTrappingAreasRMBL092005.ppt. Big Flat Rock, Gooseberry, Yearling burrow, Cantalope burrow, Steep slope, Slope, Main Talus, Talus, Cliff NorthPicnic.JPG jpg ecogrid://knb/bowdish.158.1 North_Picnic North Picnic North Picnic Marmot tagging area at RMBL trapping area NorthPicnicMain.JPG North Picnic: main area - photo includes Rocky Mountian Biological Laboratory (RMBL) Marmot trapping areas. This is part of the PowerPoint Slidshow MarmotTrappingAreasRMBL092005.ppt. Gooseberry, Yearling burrow, Cantalope, Slope, Main Talus, Steep Slope NorthPicnicMain.JPG jpg ecogrid://knb/bowdish.160.1 NorthPicnicMain North Picnic: main area North Picnic: main area - marmot trapping areas trapping areas LowerPicnic.JPG Lower Picnic - photo includes Rocky Mountian Biological Laboratory (RMBL) Marmot trapping areas. This is part of the PowerPoint Slidshow MarmotTrappingAreasRMBL092005.ppt. LowerPicnic.JPG jpg ecogrid://knb/bowdish.163.1 LowerPicnic Lower Picnic Lower Picnic trapping area trapping area UpperPicnic.JPG Upper Picnic and Lower Picnic trapping areas - photo includes Rocky Mountian Biological Laboratory (RMBL) Marmot trapping areas. This is part of the PowerPoint Slidshow MarmotTrappingAreasRMBL092005.ppt UpperPicnic.JPG jpg ecogrid://knb/bowdish.164.1 UpperLowerPicnic Upper Picnic and Lower Picnic Upper Picnic and Lower Picnic trapping areas trapping areas WinterPicnic.JPG Winter Picnic - photo includes Rocky Mountian Biological Laboratory (RMBL) Marmot trapping areas. This is part of the PowerPoint Slidshow MarmotTrappingAreasRMBL092005.ppt. WinterPicnic.JPG jpg ecogrid://knb/bowdish.165.1 WinterPicnic Winter Picnic Winter Picnic tagging areas tagging areas